Reply to Engelhardt et al.: Inclusive fitness does maintain a heritable altruism polymorphism in Tibetan ground tits [Biological Sciences]

Testing Hamilton's rule (1) requires following a rigorous methodology, as stressed by us (2) and by Engelhardt et al. (3). Here, we would like to explain again how we did so. First, Engelhardt et al. (3) question the calculation of b and c. Our responses to their points are as...

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